Also attributes such as speed and trajectory can now be modified by single increments rather than 25% increments. The Create A Superstar mode now includes pre-loaded attribute points for created superstars, while in Create A Finisher mode, a new finisher position is the 'Corner', in which finishers can be created in corner position (Cannot be used during the Royal Rumble match). Like previous iterations, the game features several storylines starting during the card leading to the Royal Rumble, a time period dubbed the Road to WrestleMania ( WrestleMania XXVI). Along with that is the new storyline in which players choose Kofi Kingston, John Morrison, Dolph Ziggler, R-Truth, or a created superstar in a quest to end The Undertaker's undefeated WrestleMania Streak. Playable storylines involve Rey Mysterio, John Cena, Christian, and Chris Jericho.

Road to WrestleMania mode has been modified to include new features such as backstage roaming, mini-quests, and further interaction with other Superstars. However, one can only get into the contendership list depending on their rivalries, alliances, match types and most importantly, win-loss ratio.

The game reacts to the players actions and depending on the goal of the player's superstar, original or created, puts them in one of, or several, the top ten contenders lists for a specific title. The game features more than 100 cutscenes to push storylines, which can revolve around alliances, rivalries and titles, forward. These cut scenes appear randomly before, during, and after the match. The game's new 'WWE Universe' builds storylines, and integrates cut scenes and rivalries between wrestlers based on the matches that are wrestled. This game was also the last to feature Batista and Shawn Michaels as non-legends or unlockables until the release of later installments of the game. In addition, new ways to exit the cell were introduced, such as barreling through the cage wall. The Hell in a Cell match was revamped with expanded cell walls, weapons underneath the ring, steel steps near the poles, and the removal of the cell door. Raw game, chairs can be thrown at the player's opponents. For the first time since the original WWE SmackDown! vs. Inside and outside the ring, ladders can now be rested onto the ring ropes and be broken in half. Tables now break differently depending on the impact placed upon them. One of the main changes in the game is the incorporation of a new physics system that allows objects to be used more realistically, such as objects involved in the Tables, Ladders, and Chairs match. The Tables, Ladders and Chairs match features new ways to use and break objects. John Cena, Randy Orton, Rey Mysterio, and Drew McIntyre in a TLC Match.

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